The team of Mosh and Thrasher first made an appearance together in June 1994 in Extreme Championship Wrestling, losing to The Bad Breed (Axl and Ian Rotten). In December 1995, they debuted as jobbers in the WWF as the Spiders, the Arachnoids, and even used their real names. On an episode of Monday Night Raw, they received a WWF Tag Team Title shot against the Smoking Gunns.
The team officially debuted on WWF television on the November 24, 1996 episode of Superstars under their Headbangers gimmick defeating Alex Porteau and Aldo Montoya. Shortly after that, they began wearing nuns' clothing and billed as the Flying Nuns, Sister Angelica and Mother Smucker. They were also humorously billed as hailing from a monastery in the Himalayas. On an episode of Shotgun Saturday Night, Brother Love showed up to manage the Nuns. After the Nuns defeated The Godwinns, Love renamed them the Sisters of Love. The gimmick was short-lived, but they kept the skirts when they returned to being The Headbangers in January 1997. They later had T-shirts with the phrase "Real men wear skirts" and occasionally wore sports bras over their shirts. The duo often carried a boombox to the ring, and on one occasion, Mosh vomited during an interview.
Throughout 1997, The Headbangers feuded with The Godwinns, as well as Doug Furnas and Phil Lafon, often finding themselves on the losing end. They faced the Legion of Doom during their surprise return match on February 24 to a double countout. At WrestleMania 13, The Headbangers won a four-way elimination match for a shot at the WWF Tag Team Championship.
The Headbangers won the vacant Tag Team Titles at Ground Zero: In Your House after Mosh pinned Owen Hart, thanks to run-in interference by Steve Austin. Mosh and Thrasher would lose the titles to the Godwinns at Badd Blood: In Your House in October. They faced the New Age Outlaws in their debut match as a team on October 20 and frequently challenged them into the following the year for the Tag Team Titles.
Both Mosh and Thrasher entered the 1998 Royal Rumble and both lasted over ten minutes before Mosh was eliminated by Kurrgan and Thrasher by Steve Austin. They feuded with The Quebecers and The Oddities throughout the year.
The Headbangers were both meant to compete in the 1999 Royal Rumble, but Thrasher suffered a knee injury and was shelved while Mosh was attacked backstage by Mabel as a way to write both men off television. While Thrasher was injured, Mosh would be repackaged as Beaver Cleavage and then Chaz, performed in singles competition and began having problems with his on-screen girlfriend, Marianna. He was arrested on an episode of Sunday Night Heat in September after Marianna claimed he hit her. After Chaz's match on October 5, Thrasher showed GTV footage of Mariana applying makeup to create the effect of wounds. This proved her accusations false, and The Headbangers subsequently reunited. They then took on a gimmick where they dressed as the opponents that they feuded with, such as the Dudley Boyz and Mean Street Posse. The duo later turned heel and began to dress in drag, which included wearing breast cones.
Mosh entered in the 2000 Royal Rumble alone and was eliminated by Rikishi; however, The Headbangers continued performing in tag team and singles matches. Both men participated in the hardcore battle royal at WrestleMania 2000 where Thrasher held the title for forty-three seconds, defeating Joey Abs. The team continued battling the likes of Too Cool and T&A until July 2000. After being together for six long years, they separated, and Mosh joined D'Lo Brown as Chaz to form Lo Down. Thrasher focused on his singles career and was taken off WWF television, and the main roster later that month.
Thrasher was released from the WWF in December 2000, and Mosh was released from the WWF in August 2001.
The duo made their first appearance as a team in WWE since 2000 on the August 30, 2016 episode of SmackDown Live, as heels and participants of the SmackDown Tag Team Championship tournament, losing against Heath Slater and Rhyno in the first round. They returned on the November 1 episode of SmackDown, losing to The Usos in a Survivor Series qualifying match. Then they were seen again on the November 15 900th episode of SmackDown, teaming with other heel tag teams.