On June 20, 2012, The Ascension (Conor O'Brian and Kenneth Cameron) made their debut on NXT, defeating CJ Parker and Mike Dalton with their double team finisher, Downcast. They would later enter a rivalry with The Usos, with the Ascension getting the better end of most of the encounters between the two teams. The Ascension also defeated another main roster team in Justin Gabriel and Tyson Kidd on the October 3, 2012 episode of NXT. However, the team was disbanded after Cameron was released from his WWE contract on November 30, 2012.
From there O'Brian worked as a singles wrestler, keeping the Ascension gimmick on his own for some time. Following O'Brian's victory over Alex Riley on the June 5, 2013 edition of NXT, Rick Victor appeared on the ramp, getting O'Brian's attention. Victor would officially accompany O'Brian to the ring as his partner in the Ascension on the July 3 edition of NXT as O'Brian defeated Andy Baker. After winning a gauntlet match to determine the number one contenders for the NXT Tag Team Championship, the Ascension defeated the team of Adrian Neville and Corey Graves on the October 2 edition of NXT to capture the titles. On November 8, the two were renamed, with Conor O'Brian renamed to Konnor and Rick Victor renamed to Viktor.
For the majority of their title reign, The Ascension would battle and dismantle local wrestlers in mere minutes with very little to no offense from their opponents. Their first major title defense was against Too Cool (Grandmaster Sexay and Scotty 2 Hotty) at NXT's first live special, NXT Arrival, where they emerged victorious. The Ascension began demanding competition in the weeks following, moving up from local wrestlers to various combinations of teams made up from contracted NXT wrestlers. While still disabling and annihilating their opponents week after week, they were confronted by a team of masked wrestlers; El Local and Kalisto, who challenged them for their tag team titles at the second live event, NXT TakeOver. Though they were given their toughest challenge to date, The Ascension successfully defended their gold after hitting Fall of Man on Local.
The Ascension had a cameo on the main roster on the September 9, 2014 episode of Main Event, defeating Los Matadores (Diego and Fernando) in a match to promote their title defense against The Lucha Dragons (Kalisto and Sin Cara) at the NXT TakeOver: Fatal 4-Way event. At the event, The Ascension lost the titles to Kalisto and Sin Cara, ending their reign a day short of a full year at 364 days. That same night, they were in an altercation with William Regal and the debuting Hideo Itami. After feuding with and assaulting Itami for several weeks, a second international wrestler, Finn Bálor, joined Itami in his fight against both Konnor and Viktor. This led to a match on December 11 at NXT TakeOver: R Evolution, where The Ascension were defeated by Bálor and Itami.
The following night on SmackDown, vignettes began to air promoting The Ascension's call-up to the main roster. They were promoted to the main roster on the December 29, 2014 episode of Raw, defeating The Miz and Damien Mizdow. On the January 2 and 9 episodes of SmackDown and the January 5 episode of Raw, The Ascension easily defeated local preliminary wrestlers in less than a minute, and also cementing their heel status after comparing themselves to prominent tag teams of the past including The Powers of Pain, The Road Warriors and Demolition.
On the January 19 episode of Raw, the Ascension confronted the reuniting New World Order members Kevin Nash, Scott Hall and X-Pac. However, the Ascension were then interrupted, attacked by and laid out by two other past-generation teams: the Acolytes Protection Agency and the New Age Outlaws. The storyline culminated in the Ascension defeating the New Age Outlaws at the Royal Rumble pay-per-view.
On the February 12, 2015 episode of SmackDown, the Ascension's main roster winning streak (since their call up) ended in a tag team turmoil (gauntlet) match; they entered against Daniel Bryan and Roman Reigns and were eliminated via disqualification. On the February 23 episode of Raw, the Ascension suffered their first pinfall loss on the main roster, courtesy of the Prime Time Players. Although they defeated The Prime Time Players in a rematch on the next week's episode of Superstars, the Ascension's fortunes then vastly declined; between early March to the end of July 2015, they appeared regularly on Main Event and Superstars, and started a losing streak. At WrestleMania 31, Konnor and Viktor made their WrestleMania debuts in the André the Giant Memorial Battle Royal, with neither of the duo winning the match. In May, The Ascension started a feud against The Meta Powers and defeated them on the Payback pre-show. At Elimination Chamber on May 31, they participated in an Elimination Chamber match for the tag titles, where they eliminated Los Matadores and the Lucha Dragons until they were eliminated by The Prime Time Players.
On the September 3 episode of SmackDown, The Ascension attacked Neville as he was making his way to the ring against Stardust, aligning themselves with Stardust in the process and thus forming a group known as "The Cosmic Wasteland", wherein Stardust called The Ascension his "henchmen". At Night of Champions, Stardust and The Ascension defeated Neville and The Lucha Dragons in a six-man tag team match on the pre-show, but were subsequently defeated in two rematches. On the Survivor Series kickoff, The Cosmic Wasteland, The Miz, and Bo Dallas were defeated by The Dudley Boyz, Neville, Goldust and Titus O'Neil in a five-on-five traditional Survivor Series elimination tag team match. In May 2016, The Cosmic Wasteland stable disbanded after Stardust left the company. At WrestleMania 32, Konnor and Viktor entered the Andre the Giant Memorial Battle Royal, both with failing efforts. On April 16, 2016, WWE suspended Konnor for sixty days for a second violation of the wellness policy. Following the suspension, Viktor began competing in singles competition, mainly on Superstars and Main Event. Konnor made his return during the June 20 WWE Superstars taping, teaming with Viktor to face The Golden Truth (Goldust and R-Truth).
On July 19 at the 2016 WWE draft, The Ascension were drafted to SmackDown brand in the eighth round. On the July 26 episode of SmackDown Live, both members competed in a WWE World Championship number one contendership six-pack qualifying battle royal, which was won by Apollo Crews. At SummerSlam, The Ascension competed in a 12-man tag team match, where their team lost. They then entered the SmackDown Tag Team Championship tournament, where they were eliminated in the first round by The Usos. On the TLC: Tables, Ladders & Chairs pre-show, The Ascension, The Vaudevillains and Curt Hawkins lost a ten-man tag team match against Apollo Crews, American Alpha and The Hype Bros. At the 2017 Elimination Chamber pay-per-view, they fought in a tag team turmoil match for the SmackDown Tag Team Championship, but were the last team to be eliminated, losing to the reigning champions American Alpha. At WrestleMania 33, they once again entered the André the Giant Memorial Battle Royal, but neither were successful. At Money in the Bank, The Ascension lost to Breezango (Tyler Breeze and Fandango) after they claimed responsibility for a vandal attack in their office. On the June 27 episode of SmackDown Live, The Ascension was interrogated by Breezango on "Fashion Vice", admitting that they didn't commit the crime and only took responsibility, because they wanted a match at the pay-per-view. In September, the duo turned face when they started trying to befriend Breezango during the latters' 'Fashion Files' segments. On the January 9, 2018 episode of SmackDown Live, The Ascension suffered a loss to The Bludgeon Brothers, for a while the duo would only appear as supporting characters in Breezango's Fashion Files skits. At WrestleMania 34, the duo were participants in the fifth annual André the Giant Memorial Battle Royal, with Konnor being the first of the duo to be eliminated followed minutes later by Viktor.
On April 16, The Ascension were traded to the Raw brand as part of Superstar Shake-up. On the April 23 episode of Raw, The Ascension, once again as heels, faced the team of Matt Hardy and Bray Wyatt in a losing effort. The two were participants in the Greatest Royal Rumble match, with Viktor being eliminated by Daniel Bryan, and Konnor being eliminated by Elias and The New Day (Kofi Kingston and Xavier Woods). On the April 12, 2019 episode of Main Event, The Ascension lost to Heavy Machinery (Otis and Tucker), in what would be their final match in WWE. On December 8, 2019, after eight months of inactivity, The Ascension were released from their WWE contracts.